Tabanomorpha

Infraorder of insects

Tabanomorpha
Tabanus sudeticus
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Suborder: Brachycera
Infraorder: Tabanomorpha
Families
  • Alinkidae—extinct (Triassic)
  • Athericidae—ibis flies
  • Austroleptidae
  • Bolbomyiidae
  • Eostratiomyiidae—extinct (Middle Jurassic)
  • Oreoleptidae
  • Pelecorhynchidae
  • Rhagionidae—snipe flies
  • Tabanidae—horse and deer flies

The Brachyceran infraorder Tabanomorpha is a small group that consists primarily of two large families, the Tabanidae (horse and deer flies) and Rhagionidae (snipe flies), and an assortment of very small affiliated families, most of which have been (or could be, or sometimes are) included within the Rhagionidae.[1]

Description

Adult Tabanomorpha typically have a convex face and antenna bearing styli. The forewing has a costa along its entire perimeter (though its posterior portion may be weaker), while the tarsi have pulvilliform empodia.[2] Males have eyes that are nearly or fully holoptic[2] and have an endoaedeagal process which is usually quite long and distinct.[3] Females have the cercus always flattened.[3]

Larval Tabanomorpha have a retractable head capsule and a brush of setae just under the fold of the integument.[3]

Ecology

Adults of most species feed on nectar and pollen, but blood-feeding (hematophagy) occurs in the majority of female Tabanidae, some Rhagionidae and an Athericidae. Blood-feeding is believed to have evolved multiple times within the group.[4]

Larvae are mostly predators in terrestrial, aquatic or semi-aquatic habitats.[4]

Classification

The infraorder Vermileonomorpha is often included within the Tabanomorpha, though the most recent classifications place them as its sister taxon. There are also some classifications that place the Nemestrinoidea within the Tabanomorpha, though this is not widely accepted. There are two superfamily-level lineages currently recognized within Tabanomorpha; the Tabanoidea and the Rhagionoidea (the latter comprising Austroleptidae, Bolbomyiidae, and Rhagionidae).[1]

References

  1. ^ a b Kerr, P.H. 2010: Phylogeny and classification of Rhagionidae, with implications for Tabanomorpha (Diptera: Brachycera). Zootaxa, 2592: 1–133.
  2. ^ a b "Infraorder Tabanomorpha". bugguide.net. Retrieved 2022-06-21.
  3. ^ a b c Zloty, Jack; Sinclair, Bradley J.; Pritchard, Gordon (2005). "Discovered in our backyard: a new genus and species of a new family from the Rocky Mountains of North America (Diptera, Tabanomorpha): XXXX". Systematic Entomology. 30 (2): 248–266. doi:10.1111/j.1365-3113.2005.00270.x. S2CID 84963789.
  4. ^ a b Wiegmann, Brian M; Tsaur, Shun-Chern; Webb, Donald W; Yeates, David K; Cassel, Brian K (2000-09-01). "Monophyly and Relationships of the Tabanomorpha (Diptera: Brachycera) Based on 28S Ribosomal Gene Sequences". Annals of the Entomological Society of America. 93 (5): 1031–1038. doi:10.1603/0013-8746(2000)093[1031:MAROTT]2.0.CO;2. ISSN 0013-8746. S2CID 86268974.
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Extant Diptera families
Suborder Nematocera
Axymyiomorpha
Culicomorpha
Culicoidea
  • Dixidae (meniscus midges)
  • Corethrellidae (frog-biting midges)
  • Chaoboridae (phantom midges)
  • Culicidae (mosquitoes)
Chironomoidea
  • Thaumaleidae (solitary midges)
  • Simuliidae (black flies)
  • Ceratopogonidae (biting midges)
  • Chironomidae (non-biting midges)
Blephariceromorpha
  • Blephariceridae (net-winged midges)
  • Deuterophlebiidae (mountain midges)
  • Nymphomyiidae
Bibionomorpha
Bibionoidea
  • Bibionidae (march flies, lovebugs)
Anisopodoidea
  • Anisopodidae (wood gnats)
Sciaroidea
(fungus gnats)
Perissommatomorpha
Psychodomorpha
Scatopsoidea
Psychodoidea
  • Psychodidae (moth flies)
Ptychopteromorpha
  • Ptychopteridae (phantom crane flies)
  • Tanyderidae (primitive crane flies)
Tipulomorpha
Trichoceroidea
  • Trichoceridae (winter crane flies)
Tipuloidea
(crane flies)
  • Cylindrotomidae (long-bodied crane flies)
  • Limoniidae (limoniid crane flies)
  • Pediciidae (hairy-eyed craneflies)
  • Tipulidae (large crane flies)
Suborder Brachycera
Asilomorpha
Asiloidea
Empidoidea
Nemestrinoidea
  • Acroceridae (small-headed flies)
  • Nemestrinidae (tangle-veined flies)
Muscomorpha
Aschiza
Platypezoidea
  • Ironomyiidae (ironic flies)
  • Lonchopteridae (spear-winged flies)
  • Opetiidae (flat-footed flies)
  • Phoridae (scuttle flies, coffin flies, humpbacked flies)
  • Platypezidae (flat-footed flies)
Syrphoidea
  • Pipunculidae (big-headed flies)
  • Syrphidae (hoverflies)
Schizophora
Acalyptratae
Conopoidea
  • Conopidae (thick-headed flies)
Tephritoidea
  • Pallopteridae (flutter flies)
  • Piophilidae (cheese flies)
  • Platystomatidae (signal flies)
  • Pyrgotidae
  • Richardiidae
  • Tephritidae (peacock flies)
  • Ulidiidae (picture-winged flies)
Nerioidea
  • Cypselosomatidae
  • Micropezidae (stilt-legged flies)
  • Neriidae (cactus flies, banana stalk flies)
Diopsoidea
Sciomyzoidea
Sphaeroceroidea
Lauxanioidea
Opomyzoidea
Ephydroidea
  • Camillidae
  • Curtonotidae (quasimodo flies)
  • Diastatidae (bog flies)
  • Drosophilidae (vinegar and fruit flies)
  • Ephydridae (shore flies)
  • Mormotomyiidae (frightful hairy fly)
Carnoidea
Lonchaeoidea
Calyptratae
Muscoidea
  • Anthomyiidae (cabbage flies)
  • Fanniidae (little house flies)
  • Muscidae (house flies, stable flies)
  • Scathophagidae (dung flies)
Oestroidea
Hippoboscoidea
  • Glossinidae (tsetse flies)
  • Hippoboscidae (louse flies)
  • Nycteribiidae (bat flies)
  • Streblidae (bat flies)
Stratiomyomorpha
Stratiomyoidea
  • Pantophthalmidae (timber flies)
  • Stratiomyidae (soldier flies)
  • Xylomyidae (wood soldier flies)
Tabanomorpha
Rhagionoidea
Tabanoidea
Vermileonomorpha
Vermileonoidea
Xylophagomorpha
Xylophagoidea
  • Xylophagidae (awl flies)
Taxon identifiers
Tabanomorpha